Africa Wild Insect Book Flies (Diptera)

Discussions and information on all Southern African Invertebrates

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ExFmem
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Re: Africa Wild Insect Book Flies: Diptera Bombyliidae

Post by ExFmem »

Bee Fly Possibly Petrorossia sp.
Family: Bombyliidae. Subfamily: Anthracinae. Tribe: Xeramoebini

Image
Kgalagadi Transfrontier Park (October)

Description
Relatively narrow wings, alula reduced and narrow; discal cell long. Crossvein r-m before middle of discal cell.
Face rounded. Hind margin of eye with a bisecting line. Body elongate, abdomen flattened, with hairs and sparse small scales.

http://www.diptera.info/forum/viewthrea ... d_id=59508


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Diptera, Brachycera, Empidoidea, Dolichopodidae

Post by Toko »

Family Dolichopodidae (Dolichopodid Flies, Long-legged Flies)

Dolichopodid Flies are minute to small in size with bright metallic colors, mostly green or bronze. They usually have slender body and large eyes. They have taped long antenna. Their legs are long so they have other common name Long-legged Flies. Their wings are clear or marked with darker areas towards the wing tips. Wing venation is characteristic. They are very good fliers but usually found resting on large green leaves. They are also found rest on tree trunks, muddy ground and river rocks.

They are predator. Adult Dolichopodid Flies' mouthparts are modified for predation, but not elongated. They feed on smaller soft body insects such as aphids. We usually found those Dolichopodid Flies resting on large leaves. They can also be found on tree trunks, mud flats and river rocks.

Larvae are maggot-like. They are usually found in moist soil and under tree bark. They are either scavengers or predators of other insects larvae.

Links: Wikipedia


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Re: Africa Wild Insect Book Flies Photos & Descriptions

Post by Klipspringer »

Bathroom Moth Fly, Drain Fly Clogmia albipunctata
Family Psychodidae. Subfamily Psychodina.

Clogmia albipunctata.jpg
Clogmia albipunctata.jpg (38.01 KiB) Viewed 1727 times
Kruger National Park © ExFmem

Psychodidae:
Adults small- to medium-sized lower Diptera (wing length: 1–6 mm), with body shape humpbacked; wings held erect, roof-like or horizontally over abdomen at rest; adults with flight short and erratic; antennae with hyaline sensory rods (termed ascoids); palpus 3- to 5-segmented; ocelli absent; wing shape variable, with vein Sc, 4–5 R veins, 4 M veins, 1–2 Cu veins of which CuP is restricted to wing base; crossveins reduced, or confined to Sc and/or basal cells. Morphology highly variable between subfamilies.

Psychodinae:
Habitus of adult stout, short-legged, vestiture developed. Head with compound eye reniform, usually with dorsal extension; antenna with 8–14 flagellomeres; ascoids paired, or multiple, highly variable in shape; palpus 4-segmented; female mouthparts without functional mandibles; males of some genera with club- or sac-shaped structures posteriorly on head. Thorax with wing lanceolate to oval, distally rounded or pointed; vein R 5-branched; jugum usually well-developed; prothorax and/or mesothorax in males of some genera with eversible, or permanently everted structures for display and pheromone release. Abdomen with male terminalia inverted by full rotation of segment 9; gonocoxite usually curved cylindrical, with parameral complex only partially sclerotised; gonostylus usually tapering, but variable; male with paired testes and 3 pairs of accessorial glands; ejaculatory duct unpaired, unsclerotised (muscular), with 1 phallotrema; epandrium present, with jointed muscular appendages present, invariably with tenacula; cercus absent; female with single genital opening; spermathecae and ducts reduced; structures superficially similar to spermatheca often present.

Clogmia Enderlein (Psychodinae).
A genus of ca 30 described species occurring in all zoogeographical regions, except Antarctica. The genus is in need of revision, but six species occur in the Afrotropics, described from Cabo Verde, Madagascar, Malawi, Sierra Leone and South Africa. The type species and probably most commonly encountered species, C. albipunctata, is a widespread synanthropic moth-fly, occurring commonly in bathrooms and kitchens in tropical, subtropical and warm-temperate climates throughout the world, including most Afrotropical countries. It was considered as an invasive
alien species in South Africa, but as its geographical origin is not known, it may well be native to the Afrotropical Region. The majority of species develop in water-filled tree-holes and other phytotelmata. Keys to species are not available.

Description
Clogmia albipunctata can reach a length of 3–5 m. Both the body and the wings are covered with thick gray-brown hair. The wings are pointed, and have some white spots. Despite being a Diptera (a fly), it has a similar appearance to that of a small moth. The antennae show dense setae, each segment with separate whorls.

Biology
The adults do not feed and live for about 12 days. They spend most of their life perched on walls. They move rarely, and with weak flight. The larvae live in aquatic environments, feeding on organic decaying matter, and take about 18 days to turn into a pupa, which develops into an adult after 5 days.
Although they are considered harmless, some cases of myiasis caused by the larvae of this insect are reported in the literature.

Distribution
Angola, Burundi, Cape Verde, D.R. Congo, Ghana, Kenya, Madagascar, Malawi, Mauritius, Nigeria, Senegal, South Africa, Tanzania, UAE, Uganda; Nearctic, Neotropical, Oceanic, Oriental, Palearctic.

Clogmia albipunctata may occur in large numbers in sewage treatment plants, hospitals, kitchens and bathrooms and is, therefore, often considered nuisance pest :O^ .


Links:
https://www.researchgate.net/publicatio ... 0/download
https://www.researchgate.net/publicatio ... c/download


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Re: Africa Wild Insect Book Flies Photos & Descriptions

Post by Klipspringer »

Black Scavenger Fly Paratoxopoda depilis = P. fasciventris
Family: Sepsidae. Subfamily: Sepsinae.

Image © ExFmem
Kruger National Park

Image

Image

The Sepsidae is a relatively small, morphologically and ecologically uniform family of thenSciomyzoidea. Most species are ant-like flies, with a narrow "waist", small or medium in size, 2–12 mm in length. The adult flies as well as their larvae are closely associated with various decaying substrates such as rotten algae, mushrooms and plants, soil that is rich in decaying organic matter, the excrement of animals and humans, and animal carcasses.

Distribution
Afrotropical: Angola, Botswana, Democratic Republic of the Congo, Ethiopia, Ivory Coast, Kenya, Malawi, Namibia, South Africa, Sierra Leone, Tanzania, Zimbabwe.

Diagnosis:
Two dorsocentral setae. Notopleuron with long dense hairs as on anepisternum. Middle tibia with one ventral seta. Hind tibia with one anteroventral seta at middle.
Male: posterolateral margin of abdominal tergite 5 with several fine hairs. Surstylus short, flattened laterally, with a small pointed projection on inner surface near base.
Female: posterior margin of abdominal tergite 3 and often anterior margin of tergite 4, orange-yellow.


Links:
http://biblio.naturalsciences.be/rbins- ... 23-144.pdf
https://sepsidnet.biodiversity.online/s ... 73-depilis


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Africa Wild Insect Book Flies - Diptera, Asilidae

Post by ExFmem »

Robber Fly Daspletis sp. possibly Daspletis hirtus Ricardo, 1925*
Family: Asilidae Subfamily: Stenopogoninae. Tribe: Enigmomorphini
*ID by Dr. Torsten Dikow (Field Museum of Natural History)


Image
Mystax entirely covering face (diagnostic to tell this genus apart from Microstylum)

Genus Daspletis Loew, 1858

Description
The following characters were extracted from Londt’s (1999) key to Afrotropical stenopogonine genera and serve as a brief generic diagnosis.
*Anatergites setose (a character effectively dividing the subfamily into two distinct groups);
*antennal post- pedicel lacking a distinct style, and tipped with a small pit enclosing a sensory spine (this feature is shared only by Microstylum Macquart, 1838);
*facial swelling occupying lower three-quarters of face, entirely setose;
*presutural dorsocentral macrosetae well developed; vein M1 not strongly arched anteriorly;
* postmetacoxal membrane long setose (these last few characters are used to separate the genus from Microstylum).

Distribution
Daspletis Loew, 1858 is a genus of stenopogonine Asilidae confined to the southern parts of the Afrotropical Region. This genus is closely related to Microstylum Macquart, 1838, and now incorporates eight southern African species.


Image
Kgalagadi Transfrontier Park, Mata Mata camp


Daspletis hirtus
Species of Daspletis can be segregated into two species groups, hermanni and vulpes. D. hirtus belongs in the latter.

Distribution
D. hirtus has been collected in the northern parts of the Northern Cape province of South Africa and at Sawmills in Zimbabwe. It will probably have a much more extensive distribution that will only be more fully appreciated when more sampling has been undertaken, especially in the generally poorly sampled central parts of Botswana. There does not appear to be any correlation with any of the better known centres of plant endemism (Van Wyk & Smith 2001).

Biology
NMSA prey records include: Diptera (1): Asilidae, Neolophonotus sp.. Hemiptera (1): Pentatomidae. Hymenoptera (1): Formicidae, Halticidae. Lepidoptera (4): Pieridae, Colotis agoye, unidentified moths (1).

Links:
https://bioone.org/journals/african-inv ... .0104.full


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Re: Africa Wild Insect Book Flies (Diptera)

Post by ExFmem »

Robber Fly Acnephalomyia sp.*
Family: Asilidae Subfamily: Stenopogoninae
*ID by Dr. Torsten Dikow (Field Museum of Natural History)


Image
Kgalagadi Transfrontier Park

New Genus Acnephalomyia

In revising the taxonomy of Acnephalum Macquart, 1838 it was discovered that the Afrotropical species are not congeneric when compared with the Palaearctic type species (A. olivierii Macquart, 1838 from Greece), which is congeneric with species of Pycnopogon Loew, 1847. As a consequence all Palaearctic Pycnopogon species are transferred to Acnephalum which takes precedence over Pycnopogon. In addition it was found that Sporadothrix Hermann, 1907, a genus previously synonymised with Acnephalum, is worthy of full generic reinstatement. The new genus Acnephalomyia is provided for Afrotropical species previously assigned to Acnephalum.

Species of Acnephalomyia:
Acnephalomyia andrenoides, Acnephalomyia dorsals, Acnephalomyia eremia, Acnephalomyia iota, Acnephalomyia leukoros, Acnephalomyia platygaster, Acnephalomyia probolos


Description (subfamily Stenopogoninae asilids that belong in the new Genus Acnephalomyia):

Head: Antennal postpedicel elongate, style composed of 3 elements (2 slender segments and terminal spine-like seta); head clearly wider than high in anterior view (not more or less circular); face slightly convex; mystax long, covering entire face; vertex distinctly excavated; angle of divergence of frons/vertex in anterior view < 20°; palpi 2- segmented, well-developed; proboscis straight.

Thorax: Dorsocentrals undifferentiated; anatergites bare; metepisternal macrosetae absent; postmetacoxal area membranous; pulvilli present, but poorly developed (c. one-third length of claws); wing with cell m3 open at margin; costal vein extends around wing margin, terminating at A1 (i.e. anal cell and alula without bordering vein); stump-vein at base of R4 commonly present, even if rudimentary.

Abdomen: Segments clearly wider than long and dorsoventrally flattened; terga commonly long setose laterally; segments 1–6 clearly visible, terminal segments much reduced and withdrawn (often almost hidden from view).

Image
KTP

Distribution

This genus is a southern African endemic, largely restricted to rather arid biomes (i.e. Desert, Fynbos, Nama-Karoo and Succulent Karoo).

Image

This means that populations are found mainly in the south-western parts of southern Africa — including parts of southern Namibia, the western parts of the Northern Cape and Western Cape Provinces, and the western parts of the Eastern Cape Province of South Africa. Species occur both in the winter rainfall and summer rainfall areas of the subcontinent.

Biology

Adult Acnephalomyia fly mainly during spring and summer, most species being found resting on the ground during October and November.

Image

The few prey records that are available suggest that Acnephalomyia is a generalist, feeding on anything of an appropriate size that it gains access to.

Image
Kgalagadi Transfrontier Park



https://bioone.org/journals/African-Inv ... .0212.full


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Re: Africa Wild Insect Book Flies (Diptera)

Post by ExFmem »

Grass Robber Fly Leptogaster sp.*
Family: Asilidae Subfamily: Leptogastrinae
* ID by Dr. Torsten Dikow (Field Museum of Natural History)

Subfamily: Leptogastrinae
Members of this subfamily are rather small, elongate, and slender, and not always recognized as robber flies on first sight. This is simply an adaptation to their different life history and habitat (grasslands and nearby vegetation) and not a reason for considering this taxon a distinct family.
There are presently 17 validly recognized genera worldwide.

Description
(1) mesoanterior angles of postpronotum extended onto pronotum forming two distinct lobes (Martin 1968);
(2) mesonotal apodeme situated between these pronotal lobes (Martin 1968);
(3) metathoracic coxae positioned anteriorly (a probable adaptation for clinging to grass stems);
(4) reduction of pulvilli;
(5) reduction of alula;
(6) 2nd abdominal sternite divided into two sclerites, which are divided by a membranous ‘fenestra’ (Martin 1968);
(7) male epandrium divided into a proximal part and a movable distad surstylus (Martin 1968);
(8) absence of larval mandibles (Martin 1968).

Habitat
grasslands and nearby vegetation

Image
Kruger Nat'l Park

Genus Leptogaster

The speciose, worldwide genus Leptogaster with some 262 species is most probably paraphyletic and is one reason why a delimitation of other genera of Leptogastrinae is difficult.
A detailed study on Leptogaster is needed to examine the generic validity and generic concepts within the Leptogastrinae.


https://asiloidflies.si.edu/sites/defau ... w_2003.pdf


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Re: Africa Wild Insect Book Flies (Diptera)

Post by ExFmem »

Robber Fly Trichardis cribrata (Loew, 1858)*
Family: Asilidae Subfamily: Laphriinae
* ID by Dr. Torsten Dikow

Genus Trichardis

Trichardis (Hermann, 1906) is primarily an Afrotropical genus. It is also recorded for the Palaearctic Region where four species, including one also found in the Afrotropics, were catalogued by Lehr (1988).

There are 25 Afrotropical species. Of the 25 Afrotropical species, eight (32%) are found only in Southern Africa (apicalis, cribrata, effrena, picta, spicata, terminalis, testacea, turneri). Nine (36 %) are known only from Eastern Africa, three of these being confined to the small Yemenese islands of Socotra and Abd el Kuri. Three (12 %) species are known only from Western Africa. Two species (8 %) are widespread between East and West Africa, one (4 %) is distributed between East and Central Africa, and another is found in both East and Southern Africa.

(While this genus had not been previously recorded from any other region, this author found an undescribed species from India (Oriental Region) and described it in this paper- see link below).

Habitat
Trichardis appears to be a genus largely restricted to grassland and savannah biomes. It is absent from tropical and subtropical areas where forests dominate, and from the winter-rainfall area of the south-western parts of South Africa, which is dominated by succulents and fynbos.

Image

Biology
Nothing is known about the immature stages of any Trichardis species and biological data relating to the adults are fragmentary. It appears that all species are normally encountered resting on the ground (on sand or peddles) in savannah or woodland biomes (Londt 1994). Only six prey records are known to me. Except for a single dipteran (Tachinidae), all prey items are hymenopterans (4 Halictidae, 1 Masaridae, 1 Pompilidae). The possible predilection for Hymenoptera, and Halictidae in particular, is interesting, but probably not significant as these insects are also commonly found resting on the ground and therefore easily accessible as prey.

(Note: while the above prey records do not include any spiders, this photo shows Trichardis cribrata with a jumping spider as prey)

Image
Kruger Nat’l Park by ExF

Trichardis cribrata (Loew, 1858)

Image
Kruger Nat’l Park by ExF

Description
Head: Dark red-brown to black, finely silver pruinose except for central parts of face. Antenna orange- brown, black setose except for a few small pale yellow setae; postpedicel not markedly clavate (L:D=3.2:1). Mystax uniformly pale yellowish. Ocellar tubercle with 2 macrosetae. Proboscis and palpi dark red-brown.

Thorax: Dark red-brown to black, silver pruinose except for some bare areas. Postpronotum medially strongly silver pruinose, laterally apruinose, mesonotum apruinose except for silver lateral and posterior margins, macrosetae yellow-white, fine setulae yellowish. Scutellum apruinose except for narrow anterior margin. Anepisternum with pale yellow posterior macroseta, extensively silver pruinose except for apruinose area anteroventrally. Proepimeron anteriorly pruinose, posteriorly apruinose; katepisternum posteriorly pruinose, anteriorly apruinose; anepisternum entirely pruinose.
Legs: Dark red-brown, tibiae orange-brown proximally, pulvilli and empodium of similar length. Hind femur dark red-brown, length:height ratio 3.4:1, ventral tubercles well-developed. Hind tibia lacking ventrodistal spur.

Wing: 4.0×1.6 mm. Costal vein moderately developed around most of wing margin, weakening along anal cell and absent from alula. Membrane extensively microtrichose—discal and r5 cells entirely microtrichose.

Abdomen: Dark red-brown with narrow brown-orange posterior margins, fine pale white setose. T2 dark red-brown, pruinose except for narrow weakly silver pruinose posterior margins laterally.

Type Locality
Loew's material came from ‘Caffraria’, a term used to cover much of the eastern part of present day Southern Africa. In Dr. Londt’s study, this was the only species of Trichardis found in Kruger NP.

Distribution
The species is a southern African endemic, distributed widely within the eastern half of the subregion (eastwards of about 23°E), but does not appear to occur along the subtropical and tropical eastern coast.

Image

Biology
Adults fly during the summer months of October and March (there is a record for July that needs verification).
(Note: my photos were taken in Sept.)

Habitat
While little information is available concerning habitat, labels suggest that the species is found mainly in Acacia savannah and woodland.







https://bioone.org/journals/african-inv ... .0210.full


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Re: Africa Wild Insect Book Flies (Diptera)

Post by ExFmem »

Robber Fly Ommatius sp.*
Family: Asilidae. Subfamily: Ommatiinae (Hardy, 1927)
* ID by Dr. Torsten Dikow

Subfamily OMMATIINAE Hardy, 1927
Type genus Ommatius Wiedemann, 1821.

Image
Kruger National Park - Parfuri Picnic Area by ExF

The subfamily Ommatiinae is a diverse taxon in the Afrotropics, comprising nine genera (13 genera globally), four of which are endemic, and 133 species. Three of the four endemic genera have between 1 - 3 species each, while Afroestricus Scarbrough has 21.

Distribution
This subfamily is global in distribution, except Antarctica.

Diagnosis:
Setae on anterior surface of antennal stylus, some median ommatidia larger than surrounding ones, post pedicel short and medially broadest, post metacoxal area entirely sclerotized

Genus Ommatius Wiedemann, 1821

Image
Kruger Nat’l Park - Parfuri Picnic Area by ExF (left=male/right=female)

Ommatius (Wiedemann, 1821), the largest and most diverse genus in the subfamily, has been the focal point of ongoing revision. It is paraphyletic with respect to Afroestricus chiastoneurus. Afroestricus has only recently been described for 20 Afrotropical species previously assigned to Ommatius. Though many areas of Africa remain poorly sampled, especially arid and semiarid areas, numerous interesting and undescribed taxa have been discovered recently.

Characteristically, they perch on the ends of twigs.

Image
Kruger Nat’l Park - Parfuri Picnic Area by ExF


Image
Kruger Nat’l Park - Parfuri Picnic Area by ExF


Diagnosis
The genus can be easily distinguished by its short, pyriform third antennal segment and a long style bearing one or two rows of short hairs throughout its length.
The inclusion of more Ommatius species in future studies will help to clarify the monophyly of this worldwide, speciose genus.



http://digitallibrary.amnh.org/bitstrea ... 1/B319.pdf
https://www.mapress.com/zootaxa/2010/f/z02540p047f.pdf


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Re: Africa Wild Insect Book Flies Photos & Descriptions

Post by Klipspringer »

Bombyliid Bee Fly Phthiria sp.
Family: Bombyliidae. Subfamily: Phthiriinae.

Image © ExFmem
Feeding on Cleome angustifolia

Image © ExFmem

Image © ExFmem

Image © ExFmem

Image © ExFmem
Auchterlonie picnic site, Kgalagadi Transfrontier Park, South Africa


The Bombyliidae (bee flies) are one of the largest families of Diptera, with over 5,000 valid species described worldwide. Their high diversity may be due to the parasitoid habit of the majority of their larvae. Adults feed on nectar and pollen, and no doubt feature prominently in angiosperm pollination syndromes, although few species have been studied in detail.
Bee flies occur on all continents except Antarctica, however their highest diversities occur in semi-arid and arid environments.
The family is one of the dominant families of Diptera in the Afrotropics, with 98 genera and 1439 species known.

Adult Bombyliidae are most active in bright sunshine and tend to settle when the sky is heavily overcast. Many members of the Bombyliinae, such as Bombomyia and Systoechus , are strong and agile fliers, and males are territorial, hovering over a marker and then darting toward any insects passing nearby to investigate them, either as potential mates or as rivals. Adult Anthracinae and Lomatiinae fly close to the ground and are often found basking on patches of bare earth. Many have disruptive colour patterns which break up the outline of the settled insect. Colour patterns in these subfamilies and some genera of Bombyliinae (notably Bombylella and Bombylisoma ) may also be sexually dimorphic, with males exhibiting patches of brightly coloured or metallic scales which appear to be used in courting females.

Adults of most genera have fully developed mouthparts adapted for feeding on flowers. Both sexes feed on nectar but females also feed on pollen and many have specialised hairs on the fore tarsi for gathering pollen.

All reared species are parasitoids or predators, almost exclusively of larvae or pupae of other insects. The vast majority of Bombyliidae are ectoparasitoid; endoparasitoids are known in only three tribes belonging to two distantly related subfamilies, the Toxophorinae (Gerontini and Systropodini) and Anthracinae (Villini).
No Bombyliidae have evolved structures to inject eggs directly into the host as is the case in many hymenopterous parasitoids. Bombyliid larvae usually exhibit hypermetamorphosis, and contact their host while it is in the larval stage. Bee fly larvae consume the host when it is in a quiescent stage such as the mature larva, prepupa or pupa.
Some bee flies, for example Toxophorinae, lay macrotype eggs on or near the host, but most - notably the large speciose subfamilies Bombylliinae and Anthracinae - lay large numbers of small eggs which are coated in sand particles before being ejected into irregularities in the soil surface. These species have a specialised ‘sand chamber’ formed by the terminal abdominal segments of the female. Their oviposition behaviour involving the collection of sand particles accounts for the greater diversity of the sand chamber Bombyliidae in arid and semi-arid areas where patches of bare soil can be found.


Phthiria is an Old World genus of currently 73 known species, 22 of which are restricted to the Afrotropical region.
Immatures have been reared from Gelechiidae and Tortricidae.


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